Álvares F. 2011. Ecologia e Conservação do lobo, Canis lupus L., no Noroeste de Portugal. PhD Thesis. Universidade de Lisboa. 

Andersen LW, Harms V, Caniglia R, Czarnomska SD, Fabbri E, Jędrzejewska B, Kluth G, Madsen AB, Nowak C, Pertoldi C, Randi E, Reinhardt I en Stronen AV (2015) Long-distance dispersal of a wolf, Canis lupus, in northwestern Europe. Mammal Research 60: 163-168. 10.1007/s13364-015-0220-6

Ansorge, H., G. Kluth en S. Hahne 2006. Feeding ecology of wolves Canis lupus returning to Germany. Acta Theriologica 51 (1): 99–106.

Age of Nature (AON). 2020. Episode 2, ‘Understanding’, vanaf minuut 31. URL: [ geraadpleegd: juni 2021].

Apollonio M, Andersen R en Putman R. 2010. European ungulates and their management in the 21st century.

Arendt, M., Cairns, K., Ballard, J. et al. 2016. Diet adaptation in dog reflects spread of prehistoric agriculture. Heredity 117, 301–306.

ARK. 2021. Dood doet leven. URL: [geraadpleegd: juni 2021].

Atkins, J.L., R.A. Long, J. Pansu, J. H. Daskin, A.B. Potter, M. E. Stalmans, C. E. Tarnita en R. M. Pringle, 2019. Cascading impacts of large-carnivore extirpation in an African ecosystem. Science 364: 

Axelsson, E., Ratnakumar, A., Arendt, ML. et al. 2013. The genomic signature of dog domestication reveals adaptation to a starch-rich diet. Nature 495, 360–364.

Backeryd, J. 2007. Wolf attacks on dogs in Scandinavia 1995–2005. Ecology Institute, Swedish University

Baillon J. 2016. Le loup en France aux vingtième siècle, recherches bibiographiques: 

Baker R.O. en Timm R.M. 2017. Coyote attacks on humans, 1970-2015: implications for reducing the risks. Human-Wildlife Interactions 11: 120-132. 

Baptist, M., T. van Hattum, S. Reinhard, M. van Buuren, B. de Rooij, X. Hu, S. van Rooij, N. Polman, S. van den Burg, G. Piet, T. Ysebaert, B. Walles, J. Veraart, W. Wamelink, B. Bregman, B. Bos en T. Selnes, 2019. A nature-based future for the Netherlands in 2120. Wageningen University and Research, Wageningen. 

Barja, I. 2009. Prey and prey-age preference by the Iberian wolf Canis lupus signatus in a multiple-prey ecosystem. Wildlife Biology 15 (2): 147–154.

Barja I., F. de Miguel en F. Bárcena. 2005. Faecal marking behaviour of Iberian wolf in different zones of their territory. Folia Zoologica -Praha- 54: 21-29. 

Bergerud, Arthur T. 1971. The population dynamics of Newfoundland caribou. Wildlife Monographs 25:

BIJ12, 2021a. Dossier Wolf. URL: [geraadpleegd juni 2021]

BIJ12, 2021b. Tussenrapportage wolf. URL: [geraadpleegd 15 juni 2021].

Bjorge RR en Gunson JR (1985) Evaluation of wolf control to reduce cattle predation in Alberta. Journal of Range Management 38: 483-487. 

Black Bear Conservation Coalition 2019. URL: [geraadpleegd juni 2021]

Blanco, J.C. en Y. Cortes 2007. Dispersal patterns, social structure and mortality of wolves living in agricultural habitats in Spain. Journal of Zoology 273: 114-124.

Blanco, J.C., Y. Cortes en E. Virgos 2005. Wolf response to two kinds of barriers in an agricultural habitat in Spain. Can. J. Zool. 83: 312–323.

Boerema L., L. Freriks. A.A. en D.B. van den Brink. 2021. De juridische bescherming van de wolf in Nederland en in een aantal andere Europese landen; een juridisch onderzoek ter ondersteuning van het opstellen van Nederlands wolvenbeleid in het licht van de uitvoering van de natuurwetgeving, Boerema en Van den Brink B.V., Houwerzijl/Element Advocaten, Best.

Boitani L, Salvatori V (eds) (2019) Piano di conservazione e gestione del lupo in Italia. Ministero dell’Ambiente, e della Tutela del Territorio e del Mare, ISPRA, Unione Zoologica ItalianaBoisseaux T., O. Stefanini-Meyrignac, C. Démolis en M. Vallance. 2019. URL: [geraadpleegd: juni 2021].

Bonnell M.A. en S.W. Breck. 2017. Using resident-based hazing programs to reduce human-coyote conflicts in urban environments. Human-Wildlife Interactions 11: 146-155. 

Bosscher F. 2021. Druk verkeer op ecoduct Petrea. URL: [geraadpleegd: juni 2021].

Boyce M.S. 1992. Population Viability Analysis. Annual Review of Ecology and Systematics 23: 481-497. 10.1146/

Bovenkerk B. 2018: Wolf staat symbool voor grotere ethische discussie. URL: [geraadpleegd: juni 2021].

Bovenkerk B. en J. Keulartz. 2021. Animals in Our Midst: The Challenges of Co-existing with Animals in the Anthropocene. Korthals M en Thompson PB, (editors): Springer. 

Brainerd SM, Andrén H, Bangs EE, Bradley EH, Fontaine JA, Hall W, Iliopoulos Y, Jimenez MD, Jozwiak EA, Liberg O, Mack CM, Meier TJ, Niemeyer CC, Pedersen HC, Sand H, Schultz RN, Smith DW, Wabakken P en Wydeven AP. 2008. The Effects of Breeder Loss on Wolves. The Journal of Wildlife Management 72: 89-98.

Breck SW, Poess rel SA en Bonnell MA. 2017. Evaluating lethal and nonlethal management options for urban coyotes. Human-Wildlife Interactions 11: 133-145. 

Breck SW, Williamson R, Niemeyer C en Shivik JA. 2002. Non-lethal radio activated guard for deterring wolf depredation in Idaho: Summary and call for research. Proceedings of the Vertebrate Pest Conference 20: 223-226. 

Briedermann L. 2009. Schwarzwild. Kosmos Verlag, Stuttgart

Browne-Nuñez C, Treves A, MacFarland D, Voyles Z en Turng C. 2015. Tolerance of wolves in Wisconsin: A mixed-methods examination of policy effects on attitudes and behavioral inclinations. Biological Conservation 189: 59-71.

Bruns A, Waltert M en Khorozyan I. 2020. The effectiveness of livestock protection measures against wolves (Canis lupus) and implications for their co-existence with humans. Global Ecology and Conservation 21: e00868.

Bunewich, A.N. 1988. Dynamics of numbers and diet of wolf in Belovevezha primeval forest. Zapovedniki Belorussii 12: 108–113. [in het Russisch]

Caniglia R, Fabbri E, Mastrogiuseppe L, Randi E, 2012. Who is who? Identification of livestock predators using forensic genetic approaches. Forensic Science International: Genetics 7, 397–404.

Caniglia R, Galaverni M, Velli E, Mattucci F, Canu A, Apollonio M, Mucci N, Scandura M en Fabbri E. 2020. A standardized approach to empirically define reliable assignment thresholds and appropriate management categories in deeply introgressed populations. Scientific Reports 10: 2862. 

Canteux A. 2019. Les loups erratiques en France au XXième siècle. URL: [geraadpleegd: juni 2021].

Capitani, C., I. Bertelli, P. Varuzza, M. Scandura en M. Apollonio 2004. A comparative analysis of wolf (Canis lupus) diet in three different Italian ecosystems. Mammalian Biology 69 (1): 1–10.

Cassidy K, Mech L, Macnulty D, Stahler D en Smith D. 2017. Sexually dimorphic aggression indicates male gray wolves specialize in pack defense against conspecific groups. Behavioural Processes 136. 

Cassidy, K.A., D.W. Smith, D.R. Stahler, D.R. MacNulthy, E.E. Stahler en M.C. Metz. 2020. Territoriality and competition between wolf packs. In: Smith et al, 2020a.

CEWOLF. 2021. Website [bezocht: Juni 2021]

Chapron G, Kaczensky P, Linnell JDC, et al. 2014. Recovery of large carnivores in Europe’s modern human-dominated landscapes. Science, 346: 1517-1519. 

Ciucci, P., Mancinelli, S., Boitani, L., Gallo, O., en Grottoli, L. 2020. Anthropogenic food subsidies hinder the ecological role of wolves: Insights for conservation of apex predators in human-modified landscapes. Global Ecology and Conservation, 21, e00841.

Ciucci, P., W. Reggioni, L. Maiorano en L. Boitani 2009. Long-distance dispersal of a rescued wolf from the Northern Apennines to the Western Alps. Journal of Wildlife Management 73(8): 1300-1306.

Ciucci P, Boitani L, Francisci F en Andreoli G. 1997. Home range, activity and movements of a wolf pack in central Italy. Journal of Zoology 243: 803-819.

Claudia Russo, Silvana Mattiello, Carlo Bibbiani, Alessandro Baglini, Paolo Bongi en Claudia Facchini 2014. Impact of Wolf (Canis Lupus) on Animal Husbandry in an Apennine Province, Italian Journal of Animal Science, 13:3, DOI: 10.4081/ijas.2014.3303 

Clevenger, A.P. en N. Waltho. 2000. Factors influencing the effectiveness of wildlife underpasses in Banff National Park, Alberta, Canada. Conservation Biology 14(1): 47-56.

Clevenger, A.P. 1998. Permeability of the Trans-Canada Highway to wildlife in Banff National Park: the importance of crossing structures and factors influencing their effectiveness. In: G.L. Evink, P. Garrett, D. Zeigler en J. Berry (eds.) Proceedings of the International Conference on Wildlife Ecology and Transportation: 109-111. FL-ER-69-98. Florida Department of Transportation, Tallahassee, USA.

Clevenger, A.P., B. Chruszcz, K. Gunson and J. Wierzchowski. 2002. Roads and wildlife in the Canadian Rocky Mountain Parks - Movements, mortality and mitigation. Final report to Parks Canada. Banff, Alberta, Canada.

Crawford HM, Fontaine JB en Calver MC. 2018. Ultrasonic deterrents reduce nuisance cat (Felis catus) activity on suburban properties. Global Ecology and Conservation 15: e00444. 10.1016/j.gecco.2018.e00444

Crismer P. (2018). Boîte à outils pour la réalisation d’un plan de gestion wallon du loup (Canis lupus lupus). TFE Bachelier en agronomie, orientation forêt et nature Haute-Ecole de la Province de Liège sous la supervision du DEMNA (Gembloux). 140 pp 

Croft, S., G. Smith, P. Acevedo en J. Vicente 2018. Wild boar in focus: Review of existing models on spatial distribution and density of wild boar and proposal for next steps. EFSA report. doi:10.2903/sp.efsa.2018.EN-1490

Cubaynes S, MacNulty DR, Stahler DR, Quimby KA, Smith DW en Coulson T. 2014. Density-dependent intraspecific aggression regulates survival in northern Yellowstone wolves (Canis lupus). Journal of Animal Ecology 83: 1344-1356.

Davis, J.M. en J.A. Stamps 2004: The Effect of Natal Experience on Habitat Preferences. TRENDS in Ecology and Evolution Vol.19 No.8.

DBBW. 2019. Wölfe in Deutschland: Statusbericht 2018/2019.…

DBBW. 2020. Wölfe in Deutschland: Statusbericht 2019/2020. URL: [geraadpleegd: juni 2021].

DBBW. 2021. National damage statistics. URL: [geraadpleegd: juni 2021].

De Waal, F. 2017. Zijn we slim genoeg om te weten hoe slim dieren zijn? Atlas contact.

Dejure 2021. URL: [geraadpleegd: juni 2021].

Deck O. 2015. „Der Kofferraumwolf” - Moderne Sagen um die Einwanderung von Großbeutegreifern. Thesis. Technische Universität München. 

DelGiudice, D.G. 1998. Surplus Killing of White-Tailed Deer by Wolves in Northcentral Minnesota, Journal of Mammalogy 79(1): 227–235

Den Ouden, J., Lammertsma, D., en Jansman, H. 2020. Effecten van hoefdieren op Natura 2000-boshabitattypen op de Veluwe. (Wageningen Environmental Research rapport; No. 3013). Wageningen Environmental Research.

Desmyter S. en L. Gijsbers. 2012. Belgian canine population and purebred study for forensics by improved mitochondrial DNA sequencing. Forensic Science International: Genetics 6: 113-120.

Donfrancesco V, Ciucci P, Salvatori V, Benson D, Andersen LW, Bassi E, Blanco JC, Boitani L, Caniglia R, Canu A, Capitani C, Chapron G, Czarnomska SD, Fabbri E, Galaverni M, Galov A, Gimenez O, Godinho R, Greco C, Hindrikson M, Huber D, Hulva P, Jedrzejewski W, Kusak J, Linnell JDC, Llaneza L, López-Bao JV, Männil P, Marucco F, Mattioli L, Milanesi P, Milleret C, Mysłajek RW, Ordiz A, Palacios V, Pedersen HC, Pertoldi C, Pilot M, Randi E, Rodríguez A, Saarma U, Sand H, Scandura M, Stronen AV, Tsingarska E en Mukherjee N (2019) Unravelling the Scientific Debate on How to Address Wolf-Dog Hybridization in Europe. Frontiers in Ecology and Evolution 7. 10.3389/fevo.2019.00175

Dickie, M., R. Serrouya, R.S. McNay en S. Boutin. 2017. Faster and farther: wolf movement on linear features and implications for hunting behaviour. Journal of Applied Ecology 54: 253–2. doi: 10.1111/1365-2664.12732

Ditmer, M.A., J.R. Fieberg, R.A. Moen, S.K. Windels, S.P. Stapleton en T.R. Harris. 2018. Moose movement rates are altered by wolf presence in two ecosystems. Ecology and Evolution 2018; 8: 9017–9033. DOI: 10.1002/ece3.4402.

Drenthen, M. 2020 Het bos heeft duizend ogen - Over wolven en de noodzaak tot intersoortelijke communicatie. Zoogdier 31-3.

Drenthen, M. 2021. Coexisting with wolves in cultural landscapes: fences as communicative devices. In: Bovenkerk B en Keulartz J. 2021. Animals in Our Midst: The Challenges of Co-existing with Animals in the Anthropocene. Korthals M en Thompson PB, (editors): Springer

DREAL Rhône-Alpes. 2021. URL: [geraadpleegd: juni 2021].

Duchamp C, Chapron G, Gimenez O, Robert A, Sarrazin F, Beudels-Jamar R en Le Maho Y. (2017). Expertise collective scientifique sur la viabilité et le devenir de la population de loups en France à long terme sous la coordination ONCFS-MNHN de: Guinot-Ghestem, M., Haffner, P., Marboutin, E., Rousset, G., Savoure-Soubelet, A., 

Dufresnes C, Remollino N, Stoffel C, Manz R, Weber J-M en Fumagalli L (2019) Two decades of non-invasive genetic monitoring of the grey wolves recolonizing the Alps support very limited dog introgression. Scientific Reports 9: 148. 10.1038/s41598-018-37331-x

Edgar JP, Appleby RG en Jones DN (2007) Efficacy of an ultrasonic device as a deterrent to dingoes (Canis lupus dingo): a preliminary investigation. Journal of Ethology 25: 209-213. 10.1007/s10164-006-0004-1

Estes, J., J. Terborgh, J. Brashares, M. Power en J. Berger, 2011. Trophic downgrading of planet Earth. Science 333, 301–306

European Council (Raad van Europa). 2014. Recommendation No. 173 (2014) on hybridization between wild gray wolves (Canis lupus) and domestic dogs (Canis lupus familiaris). URL: [geraadpleegd: juni 2021]. 

Everaert J., D. Gorissen, K. Van Den Berge, J. Gouwy, J. Mergeay, C. Geeraerts, A. Van Herzele, M.-L. Vanwanseele, B. D’hondt en K. Driesen 2018. Wolvenplan Vlaanderen. Versie 7 augustus 2018. Rapporten van het Instituut voor Natuur- en Bosonderzoek 2018 (70). Instituut voor Natuur- en Bosonderzoek, Brussel, Belgium.

Evans D en Arvela M. (2011). Assessment and reporting under Article 17 of the Habitats Directive. Explanatory Notes en Guidelines for the period 2007-2012 

Fabbri E, Miquel C, Lucchini V, Santini A, Caniglia R, Duchamp C, Weber JM, Lequette B, Marucco F, Boitani L, Fumagalli L, Taberlet P en Randi E (2007) From the Apennines to the Alps: colonization genetics of the naturally expanding Italian wolf (Canis lupus) population. Molecular Ecology 16: 1661-1671. 10.1111/j.1365-294X.2007.03262.x

Faunabeheereenheid Gelderland (FBE). 2019a. Faunabeheerplan Grote Hoefdieren FBE Gelderland 2019–2025. FBE Gelderland.

Faunabeheereenheid Gelderland (FBE). 2019b. Faunabeheerplan Deelplan Monitoringsoorten 2020-2026. Faunabeheereenheid Gelderland. FBE Gelderland.

Faunabeheereenheid Gelderland (FBE). 2021. URL: [geraadpleegd: juni 2021].

Fechter, D. en L. Storch 2014. How many wolves (Canis lupus) fit into Germany? The role of assumptions in predictive rule-based habitat models for habitat generalists. PLoS ONE 9(7): e101798.

Fernández-Gil A, Cadete Da Rocha Pereira D, Dias Ferreira Pinto SM en Di Silvestre I. (2018). Large Carnivore Management Plans of Protection: Best Practices in EU Member States. Policy Department for Citizen’s Rights and Constitutional Affairs, European Parliament. 

Fernández-Gil A, Naves J, Ordiz A, Quevedo M, Revilla E en Delibes M (2016) Conflict Misleads Large Carnivore Management and Conservation: Brown Bears and Wolves in Spain. PLoS ONE 11: e0151541. 10.1371/journal.pone.0151541

Figueiredo, A.M., A.M. Valente, T. Barros, J. Carvalho, D.A. Silva, C. Fonseca, L. Madeira de Carvalho en R.T. Torres 2020. What does the wolf eat? Assessing the diet of the endangered Iberian wolf (Canis lupus signatus) in northeast Portugal. PloS one 15 (3): e0230433.

Flagstad O, C.W. Walker, C. Vila, et al. (2003) Two centuries of the Scandinavian wolf population: patterns of genetic variability and migration during an era of dramatic decline. Molecular Ecology 12, 869-880. 

Flagstad O (2020) The predator blog (vertaald uit het Noors). URL: [geraadpleegd: Juni 2021]

Food en Agrobusiness. 2020. Vraagtekens bij herkomst Noorse wolven. URL: [bezocht: Juni 2021] 

Forslund, P. (2009). Modeling growth and inbreeding in the Scandinavian wolf population. Appendix 1:3 of Suggestions for actions to strengthen the genetic status of the wolf population. Swedish Environmental Protection Agency (Naturvårdsverket) report

Francesco, C., C. Smoglica en S. Angelucci 2020: Infectious Diseases and Wildlife Conservation Medicine: The Case of the Canine Distemper in European Wolf Population. Animals 2020, 10, 2426; doi:10.3390/ani10122426

Frijlink, J. H. (1977). Patterns of Wolf Pack Movements Prior to Kills as Read from Tracks in Algonquin Provincial Park, Ont., Canada, Bijdragen tot de Dierkunde, 47(1), 131-137. doi:

Fritts, S.H., R.O. Stephenson, R.D. Hayes en L. Boitani. 2003. Wolves and humans. In: L.D. Mech en L. Boitani (eds.) Wolves: Behavior, Ecology, and Conservation: 289–316. University of Chicago Press, Chicago, USA.

Fuller, TK and Sievert, PR, “Carnivore demography and the consequences of changes in prey availability” (2001). CARNIVORE CONSERVATION. 106.Fuller, T.K., L.D. Mech en J.F. Cochrane, 2003. Wolf Population Dynamics. In: Mech L.D. en L. Boitani 2003.

FWO 2021. URL:… [geraadpleegd: juni 2021].

Galaverni M, Caniglia R, Pagani L, Fabbri E, Boattini A en Randi E (2017) Disentangling Timing of Admixture, Patterns of Introgression, and Phenotypic Indicators in a Hybridizing Wolf Population. Molecular Biology and Evolution 34: 2324-2339. 10.1093/molbev/msx169

Gehring, T.M. 1995. Winter wolf movements in northwestern Wisconsin and east-central Minnesota: a quantitative approach. MS Thesis, University of Wisconsin, Stevens Point, USA.

Gouwy, J., Van Den Berge, K., Berlengee, F., en Mergeay, J. (2019). Wolvenspecial Oktober 2019. Roofdiernieuws, (1).

Grente O, Duchamp C, Bauduin S, Opitz T, Chamaillé-Jammes S, Drouet-Hoguet N en Gimenez O. 2020. Tirs dérogatoires de loups en France: état des connaissances et des enjeux pour la gestion des attaques aux troupeaux. Faune Sauvage, Office National de la Chasse et de la Faune Sauvage.

Grient, R. van der, en A. Kamphuis 2020: Draagvlak wolf onderzoek Nederland. Motivaction.

Grilo, C., D.J. Smith en N. Klar. 2015. Carnivores: Struggling for survival in roaded landscapes. In: R. van der Ree, D.J. Smith en C. Grilo (eds.). Handbook of Road Ecology, First Edition: 300-312. John Wiley en Sons, Ltd., West Sussex, UK.

Groot Bruinderink, G.W.T.A., D.R. Lammertsma, G.A.J.M. Jagers op Akkerhuis, W. Ozinga, A.H.P. Stumpel, J.M. Baveco, en R.W. de Waal 2009. Ex ante evaluatie van maatwerk beheer van wilde zwijnen. Alterra rapport 1944, Wageningen UR.

Groot Bruinderink, G en E. van der Grift 2015: Populatiebeheer van wilde hoefdieren: nog niet goed op orde. Vakblad Natuur bos landschap, december 2015.

Groot Bruinderink, G.W.T.A., J.M. Baveco, K. Kramer, A.T. Kuiters, D.R. Lammertsma en S. Wijdeven 1999. Dynamische interacties tussen hoefdieren en vegetatie in de Oostvaardersplassen. IBN-Rapport nr 436, Wageningen.

Groot Bruinderink, G.W.T.A.; Jansman, H.A.H.; Jacobs, M.H.; Harmsen, M. 2012. De komst van de wolf (Canis lupus) in Nederland: een ‘factfinding study’ Wageningen: Alterra Wageningen UR, (Alterra- rapport 2339) - p. 69

Groot Bruinderink, G.W.T.A. en D.R. Lammertsma 2013. Voorstel voor een wolvenplan voor Nederland, versie 2.0. Alterra Wageningen UR, (Alterra- rapport 2486)

Gula R. 2008. Wolf Depredation on Domestic Animals in the Polish Carpathian Mountains. The Journal of Wildlife Management 72: 283-289. 10.2193/2006-368

Haber GC. 1996. Biological, Conservation, and Ethical Implications of Exploiting and Controlling Wolves. Conservation Biology 10: 1068-1081. 

Harper, E.K., Paul, W.J., Mech, L.D., Weisberg, S. 2008. Effectiveness of lethal, directedwolf-depredation control in Minnesota. J. Wildlife Manage. 72, 778–784.

Harmoinen J, Von Thaden A, Aspi J. 2021. Reliable Wolf-Dog Hybrid Detection in Europe Using a Reduced SNP Panel Developed for Non-Invasively Collected Samples. Research Square preprint. 

Hawley JE, Gehring TM, Schultz RN, Rossler MT en Wydeven AP (2009) Assessment of Shock Collars as Nonlethal Management for Wolves in Wisconsin. Journal of Wildlife Management 73: 518-525. 

Hawley JE, Rossler ST, Gehring TM, Schultz RN, Callahan PA, Clark R, Cade J en Wydeven AP. 2013. Developing a new shock-collar design for safe and efficient use on wild wolves. Wildlife Society Bulletin 37: 416-422. 10.1002/wsb.234

Heck, L. en G. Raschke. 1980. Die Wildsauen: Naturgeschichte, Okologie, Hege, und Jagd. Verlag Paul Parey, Hamburg, Germany.

Hegener, M. 2019. Ons wilde Oosten 2.0. Uitgeverij Stuifzand.

Hendrikx M. 2021. Overzicht van predatie door wolf (Canis lupus) op paard in Europa. Dierenzorg, HOGent. Promoter: Decombel LM, J.

Hoban S, Bruford M, D’Urban Jackson J, Lopes-Fernandes M, Heuertz M, Hohenlohe PA, Paz-Vinas I, Sjögren-Gulve P, Segelbacher G, Vernesi C, Aitken S, Bertola LD, Bloomer P, Breed M, 
Rodríguez-Correa H, Funk WC, Grueber CE, Hunter ME, Jaffe R, Liggins L, Mergeay J, Moharrek F, O’Brien D, Ogden R, Palma-Silva C, Pierson J, Ramakrishnan U, Simo-Droissart M, Tani N, Waits L en Laikre L. 2020. Genetic diversity targets and indicators in the CBD post-2020 Global Biodiversity Framework must be improved. Biological Conservation 248: 108654.

Holzapfel M., Kindervater, J., Wagner C. en Ansorge H. 2017. Nahrungsökologie des Wolfes in Sachsen von 2001 bis 2016.

Hogberg J, Treves A, Shaw B en Naughton-Treves L. 2016. Changes in attitudes toward wolves before and after an inaugural public hunting and trapping season: early evidence from Wisconsin’s wolf range. Environmental Conservation 43: 45-55. 10.1017/S037689291500017X

Hössjer O, Olsson F, Laikre L en Ryman N. 2015. Metapopulation inbreeding dynamics, effective size and subpopulation differentiation—A general analytical approach for diploid organisms. Theoretical Population Biology 102: 40-59.

Imbert, C., R. Caniglia, E. Fabbri, P. Milanesi, E. Randi, M. Serafini, E. Torretta en A. Meriggi. 2016. Why do wolves eat livestock? Factors influencing wolf diet in northern Italy. Biological Conservation 195: 156–168.

IPO, 2019. Interprovinciaal Wolvenplan. URL: [geraadpleegd: juni 2021]

Jacobs 2019: Wil Nederland wel samenleven met de wolf? URL: [geraadpleegd: juni 2021]

Jaeger JAG, Soukup T, Madriñán LF, Schwick C en Kienast F. (2011). Landscape fragmentation in Europe. Joint EEA-FOEN report. Copenhagen, Denmark: EEA. 

Janeiro-Otero, A., T.M Newsome, L.M. Van Eeden, W.J. Ripple en C.F. Dormann 2020. Grey Wolf (Canis Lupus) Predation on Livestock in Relation to Prey Availability. Biological Conservation 243.

Jansman H.A.H. 2021. Animal Conservation in the Twenty-First Century. In: Bovenkerk B., Keulartz J. (eds) Animals in Our Midst: The Challenges of Co-existing with Animals in the Anthropocene. The International Library of Environmental, Agricultural and Food Ethics, vol 33. Springer, Cham. 

Jansman H.A.H. et al., Long distance dispersal of wolf Naya: travel speed, activity and habitat use in a human dominated landscape, The Netherlands., in prep.

Jansman, H.A.H., T. Visser en D.R. Lammertsma 2019. Beïnvloedingsregio gevestigde wolf Veluwe - Opties voor een regioafbakening. Wageningen Environmental Research.

Jansman, H.A.H. en D. Sanders 2020. Wolven in Drenthe – Uitwerking van een verhoogd risicogebied. Wageningen Environmental Research.

Jarausch A, Harms V, Kluth G, et al. 2021. How the west was won: genetic reconstruction of rapid wolf recolonization into Germany’s anthropogenic landscapes. Heredity, 

Jedrzejewski W., B. Jedrzejewska, Z. Andersone-Lilley, L. Balciauskas, P. Mannil, J. Ozolins, V.E. Sidorovich, G. Bagrade, M. Kubarsepp, A. Ornicans, S. Nowak, A. Pupila en A. Zunna, 2010. Synthesizing wolf ecology and management in Eastern Europe: similarities and contrasts with North-America. In: Musiani, M., L. Boitani en P.C. Paquet (2010). The world of wolves. New perspectives on ecology, behaviour and management, pp. 207-233. University of Calgary Press, Calgary.

Jȩdrzejewski, W., B. Jȩdrzejewski, H. Okarma, K. Schmidt, K. Zub en M. Musiani 2000. Prey selection and predation by wolves in Białowieża Primeval Forest, Poland. Journal of Mammalogy 81 (1): 197–212.

Jȩdrzejewski W, Niedziałkowska M, Nowak S en Jȩdrzejewska B (2004) Habitat variables associated with wolf (Canis lupus) distribution and abundance in northern Poland. Diversity and Distributions 10: 225-233.

Jedrzejewski W, Schmidt K, Theuerkauf J, Jędrzejewska B en Kowalczyk R (2007) Territory size of wolves Canis lupus: Linking local (Białowieża Primeval Forest, Poland) and Holarctic-scale patterns. Ecography 30: 66-76. 10.1111/j.0906-7590.2007.04826.x

Jepson, P. en C. Blythe. 2020. Rewilding – the radical new science of ecological recovery. Icon Books Ltd, Omnibus Business Centre, London.

Kaartinen, S., I. Kojola en A. Colpaert. 2005. Finnish wolves avoid roads and settlements. Ann. Zool. Fennici 42: 523-532.

Klees, D., J. van Leeuwen en E. van Norren, 2019: Monitoringplan wolf. Steunstichting VZZ, Zoogdiervereniging.

Kohn, B., Frair, J., Unger, D., Gehring, T., Shelley, D., Anderson, E., and Keenlance, P. 1999. Impacts of a highway expansion project on wolves in northwestern Wisconsin. In Proceedings of the Third International Conference on Wildlife Ecology and Transportation (FL-ER-73-99), Missoula, Montana, 13–16 September 1999. Edited by G. Evink, D. Zeigler, and P. Garrett. Florida Department of Transportation, Tallahassee. pp. 53–65.

Kohn, B.E., E.M. Anderson en R.P. Thiel. 2009. Wolves, roads, and highway Development. In: A.P. Wydeven et al. (eds.) Recovery of gray wolves in the Great Lakes Region of the United States: 217-232. DOI: 10.1007/978-0-387-85952-1_14.

Khorozyan I en Waltert M (2019) How long do anti-predator interventions remain effective? Patterns, thresholds and uncertainty. Royal Society open science 6: 190826. 10.1098/rsos.190826

Kojola, I. en J. Kuittinen 2002. Wolf attacks on dogs in Finland. Wildlife Society Bulletin 30 (2): 498-501.

Kompaniyets L en Evans MA (2017) Modeling the relationship between wolf control and cattle depredation. PLoS ONE 12: e0187264. 10.1371/journal.pone.0187264

Kramer, K., Cornelissen, P., Groot Bruinderink, G.W.T.A., Kuiters, L., Lammertsma, D., Vulink, J.Th., Van Wieren, S.E., Prins, H.H.T. 2017. Effects of weather variability and geese on population dynamics of large herbivores creating opportunities for wood-pasture cycles. A modelling approach: 109-124. In: Cornelissen, P. (ed.) 2017. Large herbivores as a driving force of woodland-grassland cycles: The mutual interactions between the population dynamics of large herbivores and vegetation development in a eutrophic wetland. PhD thesis. Wageningen University, Wageningen.

Kramer-Schadt, Stephanie, Moritz Wenzler, Pierre Gras en Felix Knauer 2020: Habitatmodellierung und Abschätzung der potenziellen Anzahl von Wolfsterritorien in Deutschland. BfN-Skripten 556, 2020. DBBW.

Kraus RHS, vonHoldt B, Cocchiararo B, Harms V, Bayerl H, Kühn R, Förster DW, Fickel J, Roos C, Nowak C (2015) A single-nucleotide polymorphism-based approach for rapid and cost-effective genetic wolf monitoring in Europe based on non-invasively collected samples. Molecular Ecology Resources 15: 295-305.

Krofel M, Černe R en Jerina K (2011) Effectiveness of wolf (Canis lupus) culling to reduce livestock depredations. Acta Silvae et Ligni 95: 11-22. 

Kruuk, H. 1972. Surplus killing by carnivores. J. Zool. 166(2):233-244.

Kübarsepp, M. en H. Valdmann 2003. Winter diet and movements of wolf (Canis lupus) in Alampedja Nature Reserve, Estonia. Acta Zoologica Lituanica 13 (1): 28– 33.

Kusak, J., Skrbinšek, A.M., and Huber, D. 2005. Home ranges, movements, and activity of wolves (Canis lupus) in the Dalmatian part of Dinarids, Croatia. Eur. J. Wildl. Res. 51(4): 254–262. doi:10.1007/s10344-005-0111-2.

Lagos, L., en Bárcena, F. 2018. Spatial variability in wolf diet and prey selection in Galicia (NW Spain). Mammal research, 63(2), 125-139.

Laikre L., F Olsson, E Jansson, O Hössjer and N Ryman. 2016. Metapopulation effective size and conservation genetic goals for the Fennoscandian wolf (Canis lupus) population. Heredity 117, 

Landa A, Gudvangen K, Swenson JE en Røskaft E (1999) Factors associated with wolverine Gulo predation on domestic sheep. Journal of Applied Ecology 36: 963-973.

Langenhof, M. R., en J. Komdeur 2018: Why and how the early-life environment affects development of coping behaviours. Behavioral Ecology and Sociobiology, Vol. 72, Iss. 3, 1-32. DOI:10.1007/s00265-018-2452-3

Lanszki, J., M. Márkus, D. Újváry, Á. Szabó en L. Szemethy 2012. Diet of wolves Canis lupus returning to Hungary. Acta Theriologica 57 (2): 189–193.

Lelieveld 2012 (VU; studentenverslag): Room for wolf comeback in the Netherlands.

Lesmerises, F., C. Dussault en M.-H. St-Laurent. 2013. Major roadwork impacts the space use behaviour of gray wolf. Landscape and Urban Planning 112: 18–25.

Liberg O, Chapron G, Wabakken P, et al. (2011) Shoot, shovel and shut up: cryptic poaching slows restoration of a large carnivore in Europe. Proc. Of the Royal Society B, 279, 1730. 

Liberg O, Suutarinen J, Akesson M, et al. 2020. Poaching-related disappearance rate of wolves in Sweden was positively related to population size and negatively to legal culling. Biological Conservation 243, 108456.

Liberg. O., A. Aronson, S.M. Brainerd, J. Karlsson, H.C. Pedersen, H. Sand en P. Wabakken, 2010. The Recolonizing Scandinavian Wolf Population: Research and Management in Two Countries. In: Musiani et al, 2010.

Linnell, J. D. C., Kovtun, E. en Rouart, I. 2021. Wolf attacks on humans: an update for 2002–2020. NINA Report 1944 Norwegian Institute for Nature Research.

Linnell, J.D.C., Løe, J., Okarma, H., Blancos, J.C., Andersone, Z., Valdmann, H., Balciauskas, L., Promberger, C., Brainerd, S., Wabakken, P., Kojola, I., Andersen, R., Liberg, O., Sand, H., Solberg, E.J., Pedersen, H.C., Boitani, L. en Breitenmoser, U. (2002) The fear of wolves: a review of wolf attacks on humans. Norwegian Institute for Nature Research Oppdragsmelding, 731, 1-65. 

Linnell JDC, Salvatori V en Boitani L. 2008. Guidelines for population level management plans for large carnivores in Europe. A Large Carnivore Initiative for Europe report prepared for the European Commission (contract 070501/2005/424162/MAR/B2). 

Lopez, B. 1978: Of wolves and men. Scribner, New York.

Louchouarn N, Santiago-Ávila F, Parsons D en Treves A (2021) Evaluating how lethal management affects poaching of Mexican wolves. Royal Society open science 8: 200330. 10.1098/rsos.200330

Louv, R. 2019: Our Wild Calling - How Connecting with Animals Can Transform Our Lives--And Save Theirs. Alonquin books.

MacNulty, D.R., D.R. Stahler, T.Coulson and D.W, Smith 2020: Essential Biology of the Wolf, Foundations and Advances. In: Smith, D.W., D.R. Stahler en D.R. MacNulty (Editor) 2020a: Yellowstone Wolves Science and Discovery in the World’s First National Park. University of Chicago Press

Mancinelli, S., en P. Ciucci 2018. Beyond home: Preliminary data on wolf extraterritorial forays and dispersal in Central Italy. Mammalian Biology 93: 51–55.

Mancinelli, S., L. Boitani en P. Ciucci 2018. Determinants of home range size and space use patterns in a protected wolf (Canis lupus) population in the central Apennines, Italy. Can. J. Zool. 96: 828–838.

Mathieu R. 2021. 26 meutes de loups suivies en 2020 dans les Alpes et Préalpes françaises. Groupe PP Alpes, édition numérique, 9 p. Groupe PP Alpes. 9 p. 

Martins, I., Krofel, M., Mota, P. G., en Álvares, F. (2020). Consumption of Carnivores by Wolves: A Worldwide Analysis of Patterns and Drivers. Diversity, 12(12), 470.

Mauget, R., R. Campan, F. Spitz, M. Dardaillon, G. Janeau en D. Pépin 1984. Synthèse des connaissances actuelles sur la biologie du sanglier, perspectives de recherche. In: Symposium international sur le sanglier. Les Colloques de l’INRA 22: 15-50.

Mech, D.L. 2020: Why are Yellowstone Wolves Important? In: Smith, D.W., D.R. Stahler en D.R. MacNulty (editors). 2020a: Yellowstone Wolves Science and Discovery in the World’s First National Park. University of Chicago Press

Mech, L.D. en R.O. Peterson, 2003. Wolf-Prey Relations. In: Mech L.M. en L. Boitani eds. (2003)

Mech, L.D. 1989. Wolf population survival in an area of high road density. American Midland Naturalist 121: 387–389.

Mech, L.D. 2017. Where can wolves live and how can we live with them? Biological Conservation 210: 310–317.

Mech, L.D. en L. Boitani (editors), 2003: Wolves: behaviour, ecology, and conservation. University of Chicago Press, Chicago.

Mech, L.D., D.W. Smith en D.R. Macnulty 2015: Wolves on the hunt – the behavior of wolves hunting wild prey. University of Chicago Press.

Mech, L.D., Fritts, S.H., Radde, G., Paul, W.J., 1988. Wolf distribution in Minnesota relative to road density. Wildl. Soc. Bull. 16, 85–88.

Melis, C., P.A. Szafrańska, B. Jędrzejewska, K. Bartoń. 2006. Biogeographical Variation in the Population Density of Wild Boar (Sus scrofa) in Western Eurasia. Journal of Biogeography, 33(5), 803-811.

Meriggi, A., A. Brangi, L. Schenone, D. Signorelli en P. Milanesi 2011. Changes of wolf (Canis lupus) diet in Italy in relation to the increase of wild ungulate abundance. Ethology, Ecology en Evolution 23 (3): 195–210.

Merrill, S.B. 2000. Details of Extensive Movements by Minnesota Wolves (Canis Lupus). The American Midland Naturalist 144 (2): 428-433.

Milanesi P, A. Meriggi en E. Merli 2012. Selection of wild ungulates by wolves Canis lupus (L. 1758) in an area of the Northern Apennines (North Italy). Ethology, Ecology en Evolution 24: 81–96.

Miller, F.L., A. Gunn en E. Broughton 1985. Surplus killing as exemplified by wolf predation on newborn caribou. Can. J. Zool. 63(2): 295-300.

Miljøstyrelsen. 2021. URL: [Geraadpleegd: juni 2021].

Mills LS en Allendorf FW (1996) The one-migrant-per-generation rule in conservation and management. Conservation Biology 10: 1509-1518. 

Ministerie van Landbouw, Natuur en Voedselkwaliteit (LNV). 2008. Kamerstuk 2007-2008, 31200 XIV, nr.160, Tweede Kamer

Meerjarenprogramma Ontsnippering (MJPO). 2018. URL: [geraadpleegd: juni 2021].Morellet N, Klein F, Solberg E en Andersen R (2011) The census and management of populations of ungulates in Europe. In: Ungulate Management in Europe: Problems and Practices. Apollonio M, Andersen R en Putman R (editors)Cambridge University Press, Cambridge. p. 106-143.

Mori, E., L. Benatti, S. Lovari en F. Ferretti 2017. What does the wild boar mean to the wolf? European Journal of Wildlife Research 63: 9.

Murray DL, Smith DW, Bangs EE, Mack C, Oakleaf JK, Fontaine J, Boyd D, Jiminez M, Niemeyer C, Meier TJ, Stahler D, Holyan J en Asher VJ (2010) Death from anthropogenic causes is partially compensatory in recovering wolf populations. Biological Conservation 143: 2514-2524.

Musiani, M., L. Boitani en P.C. Paquet (editors), 2010. The World of Wolves. New Perspectives on ecology, Behaviour and Management. University of Calgary Press, Calgary.

Mysłajek RW, Tracz M, Tracz M, Tomczak P, Szewczyk M, Niedźwiecka N en Nowak S (2018) Spatial organization in wolves Canis lupus recolonizing north-west Poland: Large territories at low population density. Mammalian Biology 92: 37-44. 10.1016/j.mambio.2018.01.006

Mysłajek, R. W., Tomczak, P., Tołkacz, K., Tracz, M., Tracz, M., en Nowak, S. 2019. The best snacks for kids: the importance of beavers Castor fiber in the diet of wolf Canis lupus welpen in north-western Poland. Ethology Ecology en Evolution, 31(6), 506-513.

Mysłajek, R., S. Nowak en E.A. van der Grift. 2013. Wolven in Polen. Vakblad Natuur Bos Landschap 10 (10): 29-31.

Mysłajek, R.W., E. Olkowska, M. Wronka-Tomulewicz en S. Nowak. 2020. Mammal use of wildlife crossing structures along a new motorway in an area recently recolonized by wolves. European Journal of Wildlife Research 66:79.

Nitze, M. 2012; Schalenwildforschung im Wolfsgebiet der Oberlausitz - Projektzeitraum 2007-2010. TU Dresden

NABU. 2015. Pferd und Wolf. Wege zur Koexistenz [online]. Naturschutzbund Deutschland.

Nores, C., L. Llaneza, and Á. Álvarez. 2008. Wild boar Sus scrofa mortality by hunting and wolf Canis lupus predation: an example in northern Spain. Wildlife Biology 14:44-51, 48.

Nowak, S., R.W. Mysłajek en B. Jędrzejewska 2005. Patterns of wolf Canis lupus predation on wild and domestic ungulates in the Western Carpathian Mountains (S Poland). Acta Theriologica 50 (2): 

Nowak, S., R.W. Mysłajek, A. Kłosioska en G. Gabryś 2011. Diet and prey selection of wolves (Canis lupus) recolonising Western and Central Poland. Mammalian Biology 76 (6): 709–715.

Nowak S, Szewczyk M, Tomczak P, Całus I, Figura M en Mysłajek RW (2021) Social and environmental factors influencing contemporary cases of wolf aggression towards people in Poland. European Journal of Wildlife Research 67: 69. 10.1007/s10344-020-01455-1

Nowak S en Mysłajek R (2017) Response of the wolf (Canis lupus linnaeus, 1758) population to various management regimes at the edge of its distribution range in western Poland, 1951-2012. Applied Ecology and Environmental Research 15: 187-203. 10.15666/aeer/1503_187203

Office Fédéral de l’Environnement (OFEV). 2020. URL:… [geraadpleegd: juni 2021].

Okarma, H. 1995. The trophic ecology of wolves and their predatory role in ungulate communities of forest ecosystems in Europe. Acta Theriologica 40: 335–386.

Okarma, H. 2000: De Wolf. Thieme.

OWAD 2021. Website [bezocht: Juni 2021]

Packer, C., Holt, R., Hudson, P., Lafferty, K. en Dobson, A. 2003. Keeping the herds healthy and alert: implications of predator control for infectious disease. Ecology Letters 6, 797–802.

Peterson, R.O. en P. Ciucci 2010. The Wolf as a Carnivore. In: L.D. Mech en L. Boitani (eds). Wolves: Behavior, ecology, and conservation: 104-130. University of Chicago Press. (2003?)

Peterson, R.O, R.L. Beschta, .. E.C. Wolf 2020: Indirect effects of carnivore restoration on vegetation. In: Smith et al, 2020a.

Pilot, M., Branicki, W., Jędrzejewski, W. et al. 2010. Phylogeographic history of grey wolves in Europe. BMC Evol Biol 10, 104

Pilot M, Greco C, vonHoldt BM, Randi E, Jędrzejewski W, Sidorovich VE, Konopiński MK, Ostrander EA en Wayne RK. 2018. Widespread, long-term admixture between grey wolves and domestic dogs across Eurasia and its implications for the conservation status of hybrids. Evolutionary Applications: n/a-n/a. 10.1111/eva.12595

Pimenta V, Barroso I, Boitani L en Beja P. 2017. Wolf predation on cattle in Portugal: Assessing the effects of husbandry systems. Biological Conservation 207: 17-26.

Pimenta V, Barroso I, Boitani L en Beja P. 2018. Risks a la carte: Modelling the occurrence and intensity of wolf predation on multiple livestock species. Biological Conservation 228: 331-342.

Plaschke, M., M. Bhardwaj, H.J. König, E. Wenz, K. Dobiáš en A.T. Ford. 2021. Green bridges in a re-colonizing landscape: Wolves (Canis lupus) in Brandenburg, Germany. Conservation Science and Practice. 2021; e00364.

Hulot N en Travert S. 2018. Plan national d’actions 2018-2023 sur le loup et les activités d’élevage.…. Ministère de la transition écologique et solidaire; Ministère de l’agriculture et de l’alimentation. 

Potiek, G., W.W. Wamelink, R. Jochem and F. van Langevelde, 2012. Potential for Grey wolf Canis lupus in the Netherlands, Effects of habitat fragmentation and climate change on the carrying capacity. Wageningen, Alterra, Alterra Report 2349. 

Randi E. 2011. Genetics and conservation of wolves Canis lupus in Europe. Mammal Review 41: 99-111.

Randi E, Hulva P, Fabbri E, Galaverni M, Galov A, Kusak J, Bigi D, Bolfíková BČ, Smetanová M en Caniglia R. 2014. Multilocus Detection of Wolf x Dog Hybridization in Italy, and Guidelines for Marker Selection. PLoS ONE 9: e86409. 10.1371/journal.pone.0086409

Raynor JL, Grainger CA en Parker DP (2021) Wolves make roadways safer, generating large economic returns to predator conservation. Proceedings of the National Academy of Sciences 118: e2023251118. 10.1073/pnas.2023251118

Ražen, N., Brugnoli, A., Castagna, C. et al. 2016. Long-distance dispersal connects Dinaric-Balkan and Alpine grey wolf (Canis lupus) populations. Eur J Wildl Res 62(1): 137-142.

Reed D, O’Grady J, Brook B, Ballou J en Frankham R (2003) Estimates of Minimum Viable Population Sizes for Vertebrates and Factors Influencing Those Estimates. Biological Conservation 113: 23-34. 10.1016/S0006-3207(02)00346-4

Reinhardt I, Kluth G, Nowak C, et al. Military training areas facilitate the recolonization of wolves in Germany. Conservation Letters. 2019;12:e12635.

Reinhardt, I. en G. Kluth 2016. Abwanderungs- und Raumnutzungsverhalteen von Wólfen (Canis lupus) in Deutschland. Natur und Landschaft 91(6): 262-271.

Reinhardt, I.,Kluth, G. 2011. Pilotstudie zur Abwanderung und zur Ausbreitung von Wölfen in Deutschland [Pilot study on the dispersal and expansion of wolves in Germany]. Unpublished Final Report, F+E Vorhaben (FKZ 806 86 080). LUPUS. Accessed////

Reinhardt I, Kaczensky P, Frank J, Knauer F en Kluth G. 2020. How to deal with bold wolves. 10.19217/skr577

Ripple, W.J. en R.L. Beschta. 2012. Large predators limit herbivore densities in northern forest ecosystems. European Journal of Wildlife Research 58: 733–742.

Rossi A, Amand B, Grandmougin B en Strosser P. 2012. Les moyens de protection des troupeaux domestiques contre le loup dans les nouveaux contextes de prédation. 

Rossler ST, Gehring TM, Schultz RN, Rossler MT, Wydeven AP en Hawley JE (2012) Shock collars as a site-aversive conditioning tool for wolves. Wildlife Society Bulletin 36: 176-184. 10.1002/wsb.93

Russo C, Mattiello S, Bibbiani C, Baglini A, Bongi P en Facchini C (2014) Impact of Wolf (Canis Lupus) on Animal Husbandry in an Apennine Province. Italian Journal of Animal Science 13: 3303. 10.4081/ijas.2014.3303

Sächsisches Staatsministerium für Umwelt und Landwirtschaft. 2009. Managementplan fur den wolf in Sachsen. Safina, C: 2015 Beyond Words: What Animals Think and Feel. Henry Holt en Company, New York.

Salvatori V, Godinho R, Braschi C, Boitani L en Ciucci P. 2019. High levels of recent wolf × dog introgressive hybridization in agricultural landscapes of central Italy. European Journal of Wildlife Research 65: 73. 10.1007/s10344-019-1313-3

Sampson L en Van Patter L (2020) Advancing best practices for aversion conditioning (humane hazing) to mitigate human-coyote conflicts in urban areas. Human-Wildlife Interactions 14: 166-183. 

Sand, H., A. Eklund, B. Zimmermann, C. Wikenros en P. Wabakken 2016. Prey selection of Scandinavian wolves: single large or several small? PloS one 11 (12): e0168062.

Sankararaman S, Mallick S, Dannemann M, Prufer K, Kelso J, Paabo S, Patterson N en Reich D (2014) The genomic landscape of Neanderthal ancestry in present-day humans. Nature advance online publication. 10.1038/nature12961

Santiago-Ávila FJ, Chappell RJ en Treves A. 2020. Liberalizing the killing of endangered wolves was associated with more disappearances of collared individuals in Wisconsin, USA. Scientific Reports 10: 13881. 10.1038/s41598-020-70837-x

Santiago-Avila FJ, Cornman AM en Treves A (2018) Killing wolves to prevent predation on livestock may protect one farm but harm neighbors. PLoS ONE 13: e0189729. 10.1371/journal.pone.0189729

Sazatornil, V., Rodriguez, A., Klaczek, M., Ahmadi, M., Alvares, F., Arthur, S., et al., 2016. The role of human-related risk in breeding site selection by wolves. Biological Conservation 201: 103–110.

Schmidt J.H., Burch J.W. en MacCluskie M.C. 2017. Effects of control on the dynamics of an adjacent protected Wolf population in Interior Alaska. Wildlife Monographs 198: 1-30; DOI: 10.1002/wmon.1026

Schockert V., Fichefet V., Licoppe A. 2020. Plan d’action pour une cohabitation équilibrée entre l’homme et le loup en Wallonie. SPWARNE

Schukken, Y.H., J.C.M. van Trijp, J.J.M. van Alphen en H. Hopster (eds.) 2019: Staat van het dier – beschouwingen en opinies over de verschuivende relatie tussen mens en dier in Nederland. RDA, Den Haag.

Scott M. Brainerd, Henrik Andrén, Edward E. Bangs, Elizabeth H. Bradley, Joseph A. Fontaine, Wayne Hall, Yorgos Iliopoulos, Michael D. Jimenez, Elizabeth A. Jozwiak, Olof Liberg, Curt M. Mack, Thomas J. Meier, Carter C. Niemeyer, Hans C. Pedersen, Håkan Sand, Ronald N. Schultz, Douglas W. Smith, Petter Wabakken, and Adrian P. Wydeven “The Effects of Breeder Loss on Wolves,” Journal of Wildlife Management 72(1), 89-98, (1 January 2008).

Shivik J (2006) Tools for the Edge: What’s New for Conserving Carnivores. Bioscience 56: 253-259. 

Shivik JA, Treves A en Callahan P. 2003. Nonlethal techniques for managing predation: Primary and secondary repellents. Conservation Biology 17: 1531-1537. 10.1111/j.1523-1739.2003.00062.x

Sidorovich V. 2017. URL: [bezocht: juni 2021].

Sidorovich V, Gouwy J en Rotenko I. (2018). Unknown Eurasian lynx Lynx lynx: New findings on the species ecology and behaviour. 

Sidorovich V. en Rotenko I. 2019. Reproduction biology in grey wolves (Canis lupus) in Belarus: common beliefs versus reality. Chatyry Chverci.

Sidorovich V.E., Stolyarov V.P., Vorobei N.N., Ivanova N.V. en Je˛drzejewska B. 2007. Litter size, sex ratio and age structure of Gray Wolves Canis lupus in relation to population fluctuations in northern Belarus. Can. J. Zool. 85: 295-300. doi:10.1139/Z07-001

Sidorovich, V. 2011. Analysis of vertebrate predator-prey community: Studies within the European Forest zone in terrains with transitional mixed forest in Belarus. Tesey, Minsk, Belarus.

Sidorovich, V., A. Schnitzler, C. Schnitzler, I. Rotenko en Y. Holikava 2017. Responses of wolf feeding habits after adverse climatic events in central-western Belarus. Mammalian Biology 83: 44–50.

Sidorovich, V.E., L.L. Tikhomirova en B. Jędrzejewska 2003. Wolf Canis lupus numbers, diet and damage to livestock in relation to hunting and ungulate abundance in northeastern Belarus during 1990–2000. Wildlife Biology 9 (4): 103–111.

Sin, T., A. Gazzola, S. Chiriac en G. Rîșnoveanu 2019. Wolf diet and prey selection in the South-Eastern Carpathian Mountains, Romania. PloS one 14 (11): e0225424.

Skogen, Ketil, Olve Krange, and Helene Figari, 2017: WOLF CONFLICTS - A Sociological Study. Berghahn books.

Skogen, KI, O. Krangen en H. Figari 2017: Wolf conflicts: a sociological study. Berghahn Books. 

Smeds L, Aspi J, Berglund J, Kojola I, Tirronen K en Ellegren H (2021) Whole-genome analyses provide no evidence for dog introgression in Fennoscandian wolf populations. Evolutionary Applications 14.

Smith D.W., K.A. Cassidy, D.R. Stahler, et al. (2020): Population Dynamics and Demography. In: Smith, D.W., D.R. Stahler en D.R. MacNulty (Editor) 2020a: Yellowstone Wolves Science and Discovery in the World’s First National Park. University of Chicago Press

Smith D, White P, Stahler D, Wydeven A en Hallac D (2016) Managing wolves in the Yellowstone area: Balancing goals across jurisdictional boundaries: Managing Wolves Around Yellowstone. Wildlife Society Bulletin 40. 10.1002/wsb.677

Smith, D.W. en G. Ferguson. 2012. Decade of the wolf – Returning the wild to Yellowstone. Revised and updated version. Lyons Press, Guilford, USA.

Smith, D.W., D.R. Stahler en D.R. MacNulty. 2020a. Yellowstone Wolves Science and Discovery in the World’s First National Park. University of Chicago Press

Smith, D.W., D.R. Stahler, R. McIntyre, E.E. Stahler en K. A. Cassidy 2020b: Wolves and Humans in Yellowstone. In: Smith, D.W., D.R. Stahler en D.R. MacNulty (editors). 2020. Yellowstone Wolves Science and Discovery in the World’s First National Park. University of Chicago Press

Smuts B. 2008. Predators on Livestock Farms: A Practical Farmers Manual. The Landmark Foundation. 

Spieth PT (1974) Gene flow and genetic differentiation. Genetics 78: 961-965. 

Stahler, D.R. et al, 2020: Ecology of Family Dynamics in Yellowstone Wolf Packs. In: Smith, D.W., D.R. Stahler en D.R. MacNulty (editors). 2020. Yellowstone Wolves Science and Discovery in the World’s First National Park. University of Chicago Press

Stahler, D.R., C.C. Wilmers, A. Tallian e.a. 2020b: Competition and coexistence among Yellowstone’s meat eaters. In: Smith, D.W., D.R. Stahler en D.R. MacNulty (editors). 2020. Yellowstone Wolves Science and Discovery in the World’s First National Park. University of Chicago Press

Stone SA, Breck SW, Timberlake J, Haswell PM, Najera F, Bean BS en Thornhill DJ (2017) Adaptive use of nonlethal strategies for minimizing wolf–sheep conflict in Idaho. Journal of Mammalogy 98: 33-44. 10.1093/jmammal/gyw188

Stronen AV, Jędrzejewska B, Pertoldi C, Demontis D, Randi E, Niedziałkowska M, et al. (2013) North-South Differentiation and a Region of High Diversity in European Wolves (Canis lupus). PLoS ONE 8(10): e76454

Stubbe, C., 2008. Der Wolf in Rusland – historische Entwicklung und Probleme. Jagd en Wild 33: 

Sunde P, Collet C, Nowak C, et al. 2021. Where have all the young wolves gone? Traffic and cryptic mortality create a wolf population sink in Denmark and northernmost Germany. COnservation Letters, e12812. 

Szewczyk M, Nowak C, Hulva P, Mergeay J, Stronen AV, Bolfikova BC, Czarnomska SD, Diserens T, Fenchuk V, Figura M, De Groot A, Haidt A, Hansen M, Jansman H, Kluth G, Kwiatkowska I, Lubińska K, Michaux J, Niedźwiecka N, Nowak S, Olsen K, Reinhardt I, Romański M, Schley L, Smith S, Špinkytė-Bačkaitienė R, Stachyra P, Stępniak K, Sunde P, Thomsen P, Zwijacz-Kozica T en Mysłajek R (2021) Genetic support for the current discrete conservation unit of the Central European wolf population. Wildlife Biology. 10.2981/wlb.00809

Szewczyk M, Nowak C, Hulva P, et al. 2020. Genetic support for the current discrete conservation unit of the Central European wolf population. Wildlife Biology, wlb.00809. 

Szewczyk, M., Nowak, S., Niedźwiecka, N. et al. 2019. Dynamic range expansion leads to establishment of a new, genetically distinct wolf population in Central Europe. Sci Rep 9, 19003. 

Tanner, E., White, A., Acevedo, P. et al. 2019. Wolves contribute to disease control in a multi-host system. Sci Rep 9, 7940.

Thurber, J.A., R.O Peterson, T.D. Drummer en S.A. Thomasma. 1994. Gray wolf response to refuge boundaries and roads in Alaska. Wildlife Society Bulletin 22: 61–68.

Tikkunen M, Kojola I (2020) Does public information about wolf (Canis lupus) movements decrease wolf attacks on hunting dogs (C. familiaris)? Nature Conservation 42: 33–49.

Tobajas J, Descalzo E, Mateo R en Ferreras P. 2020. Reducing nest predation of ground-nesting birds through conditioned food aversion. Biological Conservation 242. 10.1016/j.biocon.2020.108405

Tobajas J, Gomez-Ramirez P, Maria-Mojica P, Navas I, Garcia-Fernandez AJ, Ferreras P en Mateo R (2019) Conditioned food aversion mediated by odour cue and microencapsulated levamisole to avoid predation by canids. European Journal of Wildlife Research 65. 10.1007/s10344-019-1271-9

Tobajas J, Ruiz-Aguilera MJ, Lopez-Bao JV, Ferreras P en Mateo R. 2020. The effectiveness of conditioned aversion in wolves: Insights from experimental tests. Behavioural Processes 181. 10.1016/j.beproc.2020.104259

Treves, A., R.R. Jurewicz, L. Naughton-Treves, R.A. Rose, R.C. Willging en A.P. Wydeven 2002. Wolf depredation on domestic animals in Wisconsin, 1976-2000. Wildlife Society Bulletin: 231–241.

Trouwborst en Bastmeijer, 2013;….

Trouwborst A. 2014. Exploring the Legal Status of Wolf-Dog Hybrids and Other Dubious Animals: International and EU Law and the Wildlife Conservation Problem of Hybridization with Domestic and Alien Species. Review of European, Comparative en International Environmental Law 23: 111-124.

Trouwborst A. en F.M. Fleurke. 2019. ‘Killing wolves legally – exploring the scope for lethal wolf management under European nature conservation law’ 22(3) Journal of International Wildlife Law and Policy 231-273

Valière N, Fumagalli L, Gielly L, Miquel C, Lequette B, Poulle ML, Weber JM, Arlettaz R, Taberlet P (2003) Long-distance wolf Mamm Res recolonization of France and Switzerland inferred from noninvasive genetic sampling over a period of 10 years. Anim Conserv 6:83–92 

Van Bommel, F., D. Klees, M. La Haye en J. Thissen (2020a). Analyse probleemsituaties wolf. Rapport 2020.16 V2. De Zoogdiervereniging, Nijmegen. 

Van Bommel, F., M. Pekel, L. Linnartz en R. Dorgelo 2020b: Pilot Kuddewaakhonden in de Nederlandse Schapenhouderij: Preventie van Predatie door Wolven. ARK Natuurontwikkeling, Van Bommel Faunawerk en Canine Efficiency. 

Van de Veen, H. en R. Lardinois 1991: De Veluwe natuurlijk! Een herkansing en eerherstel voor onze natuur. Schuyt en Co.

Van Den Berge K en Gouwy J. 2021. Roofdieren in opmars - nieuwe context, nieuwe uitdagingen? Natuurfocus 20: 72-83. 

Van der Grift, E. 2018: Faunabeheer: Waar is de wetenschap? Vakblad Natuur bos landschap, november 2018.

Van der Grift, E.A., F.P.J. van Bommel en D.R. Lammertsma. 2019. The effectiveness of measures to reduce roe deervehicle collisions – A review and recommendations for a field test. Report 2936. Wageningen Environmental Research, Wageningen. 

Van Der Veken, T., Van Den Berge, K., Gouwy, J., Berlengee, F., en Schamp, K. 2021. Diet of the first settled wolves (Canis lupus) in Flanders, Belgium. Lutra (Leiden), 64(1), 45-56.

Van Eeden LM, Eklund A, Miller JRB, López-Bao JV, Chapron G, Cejtin MR, Crowther MS, Dickman CR, Frank J, Krofel M, Macdonald DW, McManus J, Meyer TK, Middleton AD, Newsome TM, Ripple WJ, Ritchie EG, Schmitz OJ, Stoner KJ, Tourani M en Treves A (2018) Carnivore conservation needs evidence-based livestock protection. PLoS Biology 16: e2005577. 10.1371/journal.pbio.2005577

van Ginkel, H.A.L., 2020. Wolves, tree logs and tree regeneration. Combined effects of downed wood and wolves on the regeneration of palatable and less palatable tree species. Dissertatie, Rijksuniversiteit Groningen, Groningen.

van Ginkel, H.A.L., C. Smit en D.P.J. Kuijper, 2019b. Behavioral response of naïve and non-naïve deer to wolf urine. PLOS ONE 14(11): e0223248

van Ginkel, H.A.L., D.P.J. Kuijper, J. Schotanus en C. Smit, 2019a. Wolves and Tree Logs: Landscape-Scale and Fine-Scale Risk Factors Interactively Influence Tree Regeneration. Ecosystems 22: 

Van Haaften, J.L. 1995: Wolven op de Veluwe...? De Veluwenaar 3e jaargang. No.2. april 1995 

van Liere D., N. Siard, P. Martens en D. Jordan 2021. Conflicts with wolves can originate from their parent packs. Animals 11, 1801. 

Varley N., R. McIntyre en J. Halfpenny 2020: The Wolf Watchers. In: Smith et al, 2020.

Von Bothmer L. (2014): Pjotr und der Wolf. JÄGER - Zeitschrift für das Jagdrevier. Hamburg: Jahr Top Special Verlag, Ausgabe Februar 

Wabakken P, Sand H, Kojola I, Zimmermann B, Arnemo JM, Pedersen HC en Liberg O (2007) Multistage, Long-Range Natal Dispersal by a Global Positioning System-Collared Scandinavian Wolf. The Journal of Wildlife Management 71: 1631-1634. doi:10.2193/2006-222

Wagner, C., M. Holzapfel, G. Kluth, I. Reinhardt en H. Ansorge 2012. Wolf (Canis lupus) feeding habits during the first eight years of its occurrence in Germany. Mammalian Biology 77 (3): 196–203.

Walker L.E., J.M. Marzluff, M.C. Metz, et al. 2018. Population responses of common ravens to reintroduced gray wolves. Ecology and Evolution 8: 11158-11168. 

Wallach A, Izhaki I, Toms J, Ripple W en Shanas U. 2015. What is an apex predator? Oikos 124. 10.1111/oik.01977

Ward AI, Pietravalle S, Cowan DP en Delahay RJ. 2008. Deterrent or dinner bell? Alteration of badger activity and feeding at baited plots using ultrasonic and water jet devices. Applied Animal Behaviour Science 115: 221-232. 10.1016/j.applanim.2008.06.004

Watters, J., D.W. Smith, D.R. Stahler en D.R. MacNulty 2020: Afterword. In Smith et al, 2020a

Wells, P. 1996. Wildlife mortality on the Canadian Pacific Railway between Field and Revelstoke, British Columbia. Unpublished report.

White, P.J., K.M. Profitt en T.O. Lemke. 2012. Changes in elk distribution and group sizes after wolf restoration. American Midland Naturalist 167: 174-187.

Whittington, J., C. Cassady St. Clair en G. Mercer. 2004. Path tortuosity and the permeability of roads and trails to wolf movement. Ecology and Society 9(1): 4. [online] URL:

Whittington, J., C. Cassady St. Clair en G. Mercer. 2005. Spatial responses of wolves to roads and trails in mountain valleys. Ecological Applications 15(2): 543–553.

Wielgus RB en Peebles KA (2014) Effects of Wolf Mortality on Livestock Depredations. PLoS ONE 9: e113505. 10.1371/journal.pone.0113505

Wild, M., Hobbs, N., Graham, M. en Miller, M. W. 2011. The role of predation in disease control: a comparison of selective and nonselective removal on prion disease dynamics in deer. Journal of Wildlife Diseases 47, 78–93.

Wolven in Nederland (WIN). 2021. URL:… [geraadpleegd: juni 2021].

Wolf AB en Akey JM (2018) Outstanding questions in the study of archaic hominin admixture. PLoS Genetics 14: e1007349. 10.1371/journal.pgen.1007349

Wydeven, A.P., D.J. Mladenoff, T.A. Sickley, B.E. Kohn, R.P. Thiel en J.L. Hansen. 2001. Road density as a factor in habitat selection by wolves and other carnivores in the Great Lakes region. Endangered Species Update 18: 110–114.

Young JK, Hammill E en Breck SW (2019) Interactions with humans shape coyote responses to hazing. Scientific Reports 9. 10.1038/s41598-019-56524-6

Zimmermann, B. et al. 2015. Predator-dependent functional response in wolves: from food limitation to surplus killing. Journal of Animal Ecology 84: 102–112.

Zlatanova, D., A. Ahmed, A. Valasseva en P. Genov 2014. Adaptive diet strategy of the wolf (Canis lupus L.) in Europe: a review. Acta Zoologica Bulgarica 66 (4): 439–452.

Zones de colonisation en

Zub K, Theuerkauf J, Jedrzejewski W, Jedrzejewska B, Schmidt K en Kowalczyk R (2003) Wolf Pack Territory Marking in the Bialowieza Primeval Forest (Poland). Behaviour 140: 635-648. 10.1163/156853903322149478

Žunna, A., J. Ozoliņ en A. Pupila 2009. Food habits of the wolf Canis lupus in Latvia based on stomach analyses. Estonian Journal of Ecology 58 (2): 141.